Journal covers


Loughman, Z.J. & B.W. Williams. 2021. Cambarus ectopistes sp. nov., a new stream-dwelling crayfish (Decapoda: Cambaridae) from the French Broad, Pigeon, and Nolichucky River watersheds in the Appalachian Mountain region of North Carolina and Tennessee, USA. Zootaxa. 5082(4): 322-340.

Williams, B.W. & A.J. Phillips. 2021. A new species of Pterodrilus (Branchiobdellida: Clitellata) from the upper Yadkin-Pee Dee River Basin of North Carolina. Journal of Natural History. 55(23-24): 1491-1498.

Day, C.C., J.H. Gilbert, P.J. Manlick, J.A. Grauer, J.N. Pauli, K.T. Scribner, B.W. Williams, & P.A. Zollner. Accepted. Evaluating the legacy of multiple introductions of American martens on spatiotemporal patterns of genetic diversity. Journal of Mammalogy.

Smith, M.M., J.H. Gilbert, E.R. Olson, K.T. Scribner, T.R. Van Deelen, J.F. Van Stappen, B.W. Williams, J.E. Woodford, & J.N. Pauli. 2021. A recovery network leads to the natural recolonization of an archipelago and a potential trailing edge refuge. Ecological Applications. 31(7): e02416.

Williams, B.W. & P.G. Weaver. 2021. Asymmetrical host switching by two groups of obligate ectosymbionts of crayfishes. Western North American Naturalist. 81(1):63-70.

Pederson, J., J.T. Carlton, C. Bastidas, A. David, S. Grady, L. Green-Gavrielidis, N. Hobbs, C. Kennedy, J. Knack, M.I. McCuller, B. O’Brien, K. Osborne, S. Pankey, T. Trott. 2021. 2019 Rapid Assessment Survey of marine bioinvasions of southern New England and New York, USA, with an overview of new records and range expansions, BioInvasions Records. 10(2): 227-237.


Erséus, C., B.W. Williams, K.M. Horn, K.M. Halanych, S.R. Santos, S.W. James, M. Creuzé des Châtelliers & F.E. Anderson. 2020. Phylogenomic analyses reveal a Paleozoic clitellate annelid radiation. Zoologica Scripta. 49(5):614-640.

Parker, E., C. Jones, P.M. Arana, N.A. Alegria, R. Sarralde, F. Gallardo, A.J. Phillips, B.W. Williams & A. Dornburg. 2020. Infestation dynamics between parasitic Antarctic fish leeches (Piscicolidae) and their crocodile icefish hosts (Channichthyidae). Polar Biology. 43(6):665-677.


Glon, M.G., B.W. Williams & Z.J. Loughman. 2019. Lacunicambarus dalyae: a new species of burrowing crayfish (Decapoda: Cambaridae) from the southeastern United States. Zootaxa. 4683(3):361-380. Open access

Phillips, A.J., A. Dornburg, K.L. Zapfe, F.E. Anderson, S.W. James, C. Erséus, E. Moriarty Lemmon, A.R. Lemmon & B.W. Williams. 2019. Phylogenomic analysis of a putative missing link sparks reinterpretation of leech evolution, Genome Biology and Evolution, 11(11):3082-3093.

Vogt, G., N.J. Dorn, M. Pfeiffer, C. Lukhaup, B.W. Williams, R. Schulz & A. Schrimpf. 2019. The dimension of biological change caused by autotriploidy: A meta-analysis with triploid crayfish Procambarus virginalis and its diploid parent Procambarus fallax. Zoologischer Anzeiger. 281:53-67.

Williams, B.W. & P.G. Weaver. 2019. Two new species of Xironogiton (Clitellata: Branchiobdellida) from western North America. Journal of Natural History 53(5-6):285-300.

Horn, K.M., B.W. Williams, C. Erséus, K.M. Halanych, S.R. Santos, M. Creuzé des Châtelliers & F.E. Anderson. 2019. Na+/K+-ATPase gene duplications in clitellate annelids are associated with freshwater colonization. Journal of Evolutionary Biology. 32(6):580-591.

McCann, L.D, M.I. McCuller, J.T. Carlton, I. Keith, J.B. Geller & G.M. Ruiz. 2019. Bryozoa (Cheilostomata, Ctenostomata, and Cyclostomata) in Galapagos Island fouling communities. Aquatic Invasions. 14(1):85-131.

Perkins, M.A., B.W. Williams & W.T. Russ. 2019. Cambarus franklini, a new crayfish (Decapoda: Cambaridae) from the Catawba River Basin in western North Carolina, USA. Zootaxa. 4568(3):520-532.  

Williams, B.W. & P.G. Weaver. 2019. Redescription and Clarification of Type Designation of Entocythere cambaria Marshall, 1903 (Ostracoda: Entocytheridae) and synonymization of Entocythere illinoisensis Hoff, 1942. Zootaxa 4565(2):261-273.  

Williams, B.W., E.M. Delekta* & Z.J. Loughman. 2019. Redescription and circumscription of the Acuminate Crayfish, Cambarus acuminatus Faxon, 1884 (Decapoda: Cambaridae). Zootaxa 4560(1):040-050.

Lukas, L.*, P.G. Weaver & B.W. Williams. 2019. Distributions of two groups of obligate crayfish ectosymbionts, branchiobdellidans and entocytherid ostracods, in New England. Northeastern Naturalist 26(1):155-167.


Phillips, A.J., B.W. Williams & A.L. Braswell. 2018. Observations of cocoon deposition, emergence, and feeding in Philobdella floridana (Verrill). Southeastern Naturalist 17(4):N90-94.

Vogt, G., C. Lukhaup, B.W. Williams, M. Pfeiffer, N.J. Dorn, R. Schulz & A. Schrimpf. 2018. Morphological characterization and DNA barcoding of the marbled crayfish and new hints to its origin. Zootaxa 4524(3):329-350.

Williams, B.W. & P.G. Weaver. 2018. A historical review of the taxonomy and classification of Entocytheridae (Ostracoda: Podocopida). Zootaxa 4448(1). [Holy smokes, it’s our first monograph!]

Weaver, P.G., B.W. Williams & E.M. Sadorf. 2018. A New Record of the Sea Urchin, Rhynobrissus cuneus (Spatangoida: Brissidae) from Buxton Beach, Dare County, North Carolina, U.S.A. Southeastern Naturalist 17(3):34-39.

McCuller M.I. & J.T. Carlton. 2018. Transoceanic rafting of Bryozoa (Cyclostomata, Cheilostomata, and Ctenostomata) across the North Pacific Ocean on Japanese tsunami marine debris. Aquatic Invasions 13(1):137–162.

McCuller M.I., J.T. Carlton & J.B. Geller. 2018. Bugula tsunamiensis n. sp. (Bryozoa, Cheilostomata, Bugulidae) from Japanese tsunami marine debris landed in the Hawaiian Archipelago and the Pacific Coast of the USA. Aquatic Invasions 13(1):163–171.

Carlton J.T., J.W. Chapman, J.B. Geller, J.A. Miller, G.M. Ruiz, D.A. Carlton, M.I. McCuller, N.C. Treneman, B.P. Steves, R.A. Breitenstein, R. Lewis, D. Bilderback, D. Bilderback, T. Haga & L.H. Harris. 2018. Ecological and biological studies of ocean rafting: Japanese tsunami marine debris in North America and the Hawaiian Islands. Aquatic Invasions 13(1):1–9.

Loughman, Z.J. & B.W. Williams. 2018. Cambarus polypilosus, a new species of stream-dwelling crayfish (Decapoda: Cambaridae) from the Western Highland Rim of Tennessee, USA. Zootaxa 4403(1):171-185.

Taheri, S., S. James, V. Roy, T. Decaëns, B.W. Williams, F. Anderson, R. Rougerie, C.-H. Chang, G. Brown, L. Cunha, D.W.G. Stanton, E. Da Silva, J-H. Chen, A.R. Lemmon, E. Moriarty Lemmon, M. Bartz, D. Baretta, I. Barois, E. Lapied, M. Coulis & L. Dupont. 2018. Complex taxonomy of the ‘brush tail’ peregrine earthworm Pontoscolex corethrurus. Molecular Phylogenetics and Evolution 124: 60-70.

Larson, E.R., R.M. Egly & B.W. Williams. 2018. New records of the non-native virile crayfish Faxonius virilis (Hagen, 1870) from the upper Snake River drainage and northern Bonneville Basin of the western United States. BioInvasions Records 7.


Carlton J.T., J.W. Chapman, J.B. Geller, J.A. Miller, D.A. Carlton, M.I. McCuller, N.C. Treneman, B.P. Steves & G.M. Ruiz (2017) Tsunami-driven rafting: Transoceanic species dispersal and implications for marine biogeography. Science 357(6358): 1402–1406.

Anderson F.E., B.W. Williams, K.M. Horn, C. Erséus, K.M. Halanych, S.R. Santos & S.W. James. 2017. Phylogenomic analyses of Crassiclitellata support major northern and southern hemisphere clades and a Pangaean origin for earthworms. BMC Evolutionary Biology 17:123. [See erratum, 17:204]

Rogers, C.D., S.T. Ahyong, C.B. Boyko, C. d’ Udekem d’Acoz, et al. 2017. Images are not and should not ever be type specimens: a rebuttal to Garraffoni & Freitas. Zootaxa 4269:455-459.

Weaver, P.G.B.W. Williams. 2017. A new genus and species of entocytherid ostracod (Ostracoda: Entocytheridae) from the John Day River Basin of Oregon, U.S.A., with a key to genera of the subfamily Entocytherinae. Zootaxa 4273(4):576-586.


Shelton, K.*, P.G. Weaver & B.W. Williams. 2016. New records of entocytherid ostracods from New York and New Jersey. Northeastern Naturalist 23:525-531.  [Keri got the cover, woohoo!]

Weaver, P.G.B.W. Williams. 2016. Observations of false mating behavior in entocytherid ostracods from the northwestern United States. Invertebrate Biology 135:252-258.

Larson, E.R., M. Castellin, B.W. Williams, J.D. Olden & C.L. Abbott. 2016. Phylogenetic species delimitation for crayfishes of the genus Pacifastacus. PeerJ 4:e1915.

Howell, P.E., E.L. Koen, B.W. Williams, G.J. Roloff & K.T. Scribner. 2016. Contiguity of landscape features pose barriers to gene flow among American marten (Martes americana) genetic clusters in the Upper Peninsula of Michigan. Landscape Ecology 31:1051.

Gelder, S.R. & B.W. Williams. 2016. Global overview of Branchiobdellida (Annelida: Clitellata). Pp. 628-653. In Kawai, T., Faulkes, Z., Scholtz, G. (eds), Freshwater Crayfish: Global Overview.

Larson, E.R. & B.W. Williams. 2016. Historical biogeography of Pacifastacus crayfishes and their branchiobdellidan and entocytherid ectosymbionts in western North America. Pp. 404-447. In Kawai, T., Faulkes, Z., Scholtz, G. (eds), Freshwater Crayfish: Global Overview.


Gelder, S.R. & B.W. Williams. 2015. Branchiobdellida – Chapter 22. Pp. 551-563. In Thorp, J.H., Rogers, D.C., Tockner, K. (eds), Thorp and Covich’s Freshwater Invertebrates, 4th ed., Vol. I: Ecology and General Biology.


Keenan, S.W., M.L. Niemiller & B.W. Williams. 2014. Observations of Cambarus bartonii cavatus (Decapoda: Cambaridae) and ectosymbiotic branchiobdellidans (Annelida: Clitellata) in Cruze Cave, Knox County, Tennessee, USA. Speleobiology Notes. 6:55-61.


Williams, B.W., S.R. Gelder, H.C. Proctor & D.W. Coltman. 2013. Molecular phylogeny of North American Branchiobdellida (Annelida: Clitellata). Molecular Phylogenetics and Evolution 66:30-42.

Skelton, J., K.J. Farrell, R.P. Creed, B.W. Williams, C. Ames, B.S. Helms, J. Stoeckel & B.L. Brown. 2013. Servants, scoundrels, and hitchhikers: current understanding of the complex interactions between crayfish and their ectosymbiotic worms (Branchiobdellida). Freshwater Science 32:1345-1357.

DeWitt, P.D., B.W. Williams, Z.-Q. Lu, A.N. Fard, & S.R. Gelder. 2013. Effects of environmental and host physical characteristics on an aquatic symbiont. Limnologica 43:151-156.


Williams, B.W., H.C. Proctor & T. Clayton. 2011. Range extension of the northern crayfish, Orconectes virilis (Decapoda, Cambaridae), in the western Prairie Provinces of Canada. Crustaceana 84:451-460.

Williams, B.W., D.R. Etter, P.D. DeWitt, K.T. Scribner & P.D. Friedrich. 2011. Uncertainty in determination of sex from harvested bobcats. Journal of Wildlife Management 75:1508-1512.

Williams, B.W., K.L. Williams, S.R. Gelder & H.C. Proctor. 2011. Distribution of Entocytheridae (Crustacea: Ostracoda) in the northern Prairies of North America and reports of opportunistic clitellate annelids on crayfish hosts. Western North American Naturalist 71:276-282.

Gelder, S.R. & B.W.Williams. 2011. First distributional study of Branchiobdellida (Annelida: Clitellata) in the Great Smoky Mountains National Park with a redescription of Cambarincola holostomus Hoffman, 1963. Southeastern Naturalist 10:211-220.

Williams, B.W. & S.R. Gelder. 2011. A re-description of Cambarincola bobbi Holt, 1988, a description of a new species of Pterodrilus, and observations of sympatric species of crayfish worms (Annelida: Clitellata: Branchiobdellida) from the Cumberland River watershed in Tennessee. Southeastern Naturalist 10:199-210.

Belant, J.L., D.R. Etter, P.D. Friedrich, M.K. Cosgrove, B.W.Williams & K.T. Scribner. 2011. Comparison of techniques for sex determination of American martens. Journal of Wildlife Management 75:256-260.


Williams, B.W., C.S. Davis, & D.W. Coltman. 2010. Isolation and characterization of nine polymorphic microsatellite loci in the northern crayfish (Orconectes virilis). Conservation Genetics Resources 2:235-237.

Williams, B.W. & K.T. Scribner. 2010. Effects of multiple founder populations on spatial genetic structure of reintroduced American martens. Molecular Ecology 19:227-240.


Williams, B.W., S.R. Gelder & H. Proctor. 2009. Distribution and first report of Branchiobdellida (Annelida: Clitellata) from the Prairie Provinces of Canada. Western North American Naturalist 69:119-124.

Williams, B.W., D.R. Etter, D.W. Linden, K.F. Millenbah, S.R. Winterstein & K.T. Scribner. 2009. Noninvasive hair sampling and genetic tagging of co-distributed fishers and American martens. Journal of Wildlife Management 73:26-34.


Williams, B.W. & K.T. Scribner. 2007. Demographic and genetic evaluation of an American marten reintroduction: a comment on Swanson et al. Journal of Mammalogy 88:1342-1345.

Williams, B.W. J.H. Gilbert, P.A. Zollner. 2007. Historical perspective on the reintroduction of the fisher and American marten in Michigan and Wisconsin. USDA Forest Service, Northern Research Station, Newtown Square, PA, General Technical Report NRS-5.


Scribner, K.T., J.A. Blanchong, D.J. Bruggeman, B.K. Epperson, C.-Y. Lee, Y.-W. Pan, R.I. Shorey, B.W. Williams, H.H. Prince, S.R. Winterstein & D.R. Luukkonen. 2005. Geographical genetics: conceptual foundations and empirical applications of spatial genetic data in wildlife management.  Journal of Wildlife Management 69:1434-1453. (For correct citation see Erratum in JWM 70:614.)

Purple text = NMI Unit student interns, technicians, or volunteers (asterisked for emphasis!)

Orange text = Current or Former NC Museum of Natural Sciences staff collaborators